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??? ?????????? ??????? ?????? ?????? ???????? ?? ??????? ?????? ????????? ???????? ??????? ???????? ??????? ???????? ???????? ?????? ?? ??????? ???? ?????? ?? Saccharomyces cerevisiae ?? ???? ??????? ?????? ?? ???? Angel Yeast Co., LTD. ?? ????? ?? ????? ??? ?????? ?????? ?? ?????? ???????? ???????.

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????? 2 ?????????? ???????? ??????? ?? ????? ??????
???? ????? ?????????? ?????????? ??????????? ?????????? ?????? ??????????? ??????? ?? ?????? 1 ??????? 2.
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???????? ???????? ???????? ????????? ??????? ?? ????? ?????? ???? "????? ?????? ?????? ?????". ???? ????? "????? ????????" ???? ????? ??? ???? ???????? ??????? ?? ???? ???? ???????? ????? ????? ?? ???? ???? ????????? ?????????. ????? ???? ???????? ??????? ?? ???? ???? ??????? ???? ????? ?? ???? ?????? ??????? ????? ????? ?? ???????? ??? ?? ???? ?? ????? 3. ???? ?? ???? ?????? ???????? ???????? ?? ???? ?????? ??????? ?????? ?????? ???????? ???????? ?????? (SAR) ????????? ??????? (ISR)? ????? ????? ?? ?????? ???????? ??? ??? ???????????? ???? ??????????? ?????? ?????? ??????? ?????? ???????? ?????? ???????? ??????? ???????? ??? ?????? ?????? ???????? ?????? ????.

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???? ??? "??????? ???????" ??? ??????? ??????? ??????????????? ???????? ?? ???????. ????? ??????? ?????? ??? ?????? ??? ??? ????? ???????? ?????? ????? ??? ???? ??? ????? ?????? ?? ????? ?????? ?????? ?????? ?????? ?????? ?? ??? ???????. ??? ??????? ??? ????????? ???? ??????? ??????? ??????? ?????? ????? ????? ?????????? ? ??????????? ? ???????????? (?????????? ? ? ? ??? ????? ??????? ?? ?????????? ?? ??????????? ??????? ???????? ??????? ??? ???? ?????????? ? ????? ??????? ????? ???? ?????????? ? ????? ?????? ?????????)? ??? ???? ???? ???????? ??? ?????? ?????? ??????? ????? ????? ??????? ??????.
???? ??????? ??????? ??????? ?????? ????? ????? ????? RuBP ?????????? (1,5-????? ????? ??????? ??????????)? ?????? ????? ???? ????? ??????? (CO?) ?? 1,5-????? ????? ??????? ?????? ?????? ?? 3-???????????? ????? ??? ???? ???? ???????? ??? ?????? CO? ???? ??????? ??????? ????? ?? ????? ?????? ???????????? ?? ????????.
????? ?? "??????? ???????" ?????? ??? ??????? ???????? ????? ???????. ????? ??????? ???????? ?? ?????????? ??????? ?? ??? ???? ???? ?? ??????? ????? ?????? ???? ??? ??? ??????? ???????? ???????? ??? ????? 60? ???????? ???????? ????? ?????? 20?. ?? ???? ???????? ?????? ????????? ??????? ???????? ???????? ????? ?????? ????? ?????????? ????? ?? ?????? ????? ????? ?????? ??? ?????? ????????? ????? ???? ?? ???? ??? ????? ????????.

?????. 4 ??????? ?????? ??????? ?????? ???????? ???????
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?????. 5 ????? ??????? ???????? ????? ?? ?????????? ??????? ?? ????? ??????
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????? ??????? ???????? ????? ?? ?????????? ??????? "????? ?????" ???? "????? ?????" ??? ??????? ???????. ????? ??????? ??????? ???? ??????? ?????? ??????? ?????? ??????? ??????? ??? ??? ??????? ???? ??????? ???? ?????????. ???? ??????? ??????? ????? ????? ??????? ???????? ?? ???? ????? ?????? ?? ???? ??????? ???????? ?????? ????????? ?????? ???????? ??? ??????? ???????? ??? ???? ?? ?????? ???????? ????????? ???????? ???? ??? ???????? ???????. ??? ????? ????? ???? ???????? ????? ?????? ??????? ??????? ?????????? ????? ??? ???? ????? ?????? ????? ???? ??????? ???????? ????????.
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1. ????? ??? ?????? ????????. ???? ????????????? ??? ????? ??? ??????? ?????? ??? ?????? ????????? ?????? ????? ???? ????? ???? ???? ????? ??????? ??????? ???? ???.
2. ????? ?????? ???? ????????. ???? ?????? ???????? ???????? ?? ???? ???? ??????? ??? ????? ?????? ?????? ???????? ???????? ???????? (SAR) ????????? ???????? ???????? (ISR) ?? ?????? ?????? ??????? ???? ??????.
3. ????? ????? ??????? ??????? ?????? RuBP? ??? ????? ????? ?? ????? ??????? ?????? ????????? ?????? ????? ???? ????? ??????? ??????? ??? ??????????. ???? ??? ???? ??????? ??????? ?????? ??? ????? ????? ?????????? ?? ? ???????????? ?? ????????? ?????? ???? ??????? ????????.
4. ????? ?????? ?????? ???????? ?????? ????? ??? ?????? ????? ????? ??? ???? ??????????? ??? ?? ??? ?????? ?????? ?? ??????? ???????? ?????? ???? ???? ???????????? ?????????? ???????? ????? ?????? ?????? ??? ????????.
5. ????? ????? ??????? ???????? ??????? ????????? ?????? ??????? ??????? ???????? ?????? ???????.
?????? ?????? ??? ????????? ???????
?? ???????: 3 ????/ ?? (????? ???????)? 5 ????/ ?? (???? ???????? ???? ??????)
???? ??????: 0.3 ???/ ?? (????? ???????)? 0.5 ???/ ?? (??????? ???????? ????? ??????)
??????: ?? ?? ???? ???? ????? ?????? ?????? ????? 667 ??? ????.
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???? ??????? 1 : ???????
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????? ????????: ????? ??????? ??????? 50 ??? ?? ???????

???? ?? ???? ?????????? ?????? ??? ?????? ???????? ?????.
??? ?????? ????? ?????? ?????? ????? ???????? ?????? III ???? ????? 58% ?????? ????????? ??????? ???? ????? ?????? ??? ????? ???? ????? 66.5% ?????? ????????? ??????? ??? ??? ????? ??? ?????? ?????? 0.13 ??? ??? ???? ??? ?? ????? III ???? ?? ???? ??? ???? ????? ??????? ???? ???.
???? ??????? 2 : ?????? ??????? ?????????
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????? ????????: ????? ??????? ??????? 10 ??? ?? ??????? 20 ??? ?? ??????? 50 ??? ?? ???????

?? ?? ?????? ??????? ????????? ????? ?????? ????? ??????? ????? ??? ????? 60% ?? ??????? ??? ???? ?????? ??????? ????? ????? ?????? ????? ?????? (??? ????? ?? ??? ?????) ??? ????? 40% ?? ??????? ??? ???? ?????? ??????.
??? ????? 20 ????? ?? ???????? ????? ?????? ??????? ??????? ?????? ??? 72% ?80% ?? ????? ??????? ??? ???? ?????? ????? ????? ????? ???????? ?????? ??? 82% ?? ??? ??? ???? ?????? ??? ??? ??? ????? ?????? ???? ?? ????? ?? ?? ???? ?????? ??? ???? ??? ?? ?????????? ?? ????? ?????? ?? ???? ?? ????? ?????? ??? ??? ???? ????? ?? ?? ??????? ?????? ???????. ????? ????????: ?????
?????? ??????? 3 : ????? ????? ?????? ???? ??????? ???????
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????? ????????: ??????? 50 ??? ?? ???????

??????? ???????? ??????? ?????? ??? ??????? ??????? ??????? ?? ?????????? ??????? ??? ????? ??? ??????? ?????? ?????? ?????? ????? ????? 4.1% ?13.5% ??? ???????? ??? ??? ?????? NPK ????? 7.04%? ?6.2%? ?38.4% ??? ???????.
?????? ??????? 4 : ????? ????? ??????? ??????
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????? ????????: ?? 100 ??? ?? ???????

??? ?? 100 ??? ?? ??????? ??? ??? ????? ????? ??????? ???? ???? ?????????? ??????? III ?? ???? ?? ????? ?????? ?????? ?????? ????? ????????? ????? ????? ?????? ??????? ????? ????? ??????????? ????? ???? SPAD ????? 21.15% ?????? ??????? ??? ???? ????? ??????? ??????.